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Would Domain Keys actually help since the servers are rejecting upon connect?
There appears to have been an incident involving capacity issues within our delivery infrastructure.
The error message "451 Message temporarily deferred - 4.16.50" indicates that our MTAs are currently experiencing heavy, unusual traffic.
You may retry sending at a later time when you see this message.
However please note that emails from the mail server(s) you are using may also have recently become deprioritized due to potential issues with its mailings.
These deprioritizations were temporary but may be re-triggered if the sending IP profile continues to be poor.